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During their time on Earth, 2B and 9S suppress hostile Machine Lifeforms, coming into contact with other characters who exist on the world.

YoRHa models 2B and 9S are sent down as part of a reconnaissance force to investigate local Machine behaviour and aid local Resistance forces. The latest addition to the android forces is YoRHa, an elite android fighting force based in the orbital Bunker and communicating directly with the Council of Humanity on the Moon. Androids, created by humans, fight against the Machine Lifeforms, with the conflict turning into a prolonged proxy war. Earth has been occupied by aliens, who fight using an army of Machine Lifeforms. Nier: Automata Ver1.1a is set in the year 11945 on a post-apocalyptic Earth abandoned by humanity. Its broadcast has been twice delayed due to the COVID-19 pandemic impacting production. The series was announced in a 5th anniversary livestream, and began broadcasting in January 2023. The game's original Japanese and English cast reprised their roles. The opening and ending themes were provided respectively by Aimer and Amazarashi. Monaca, who worked on the game's music, returned to both arrange earlier pieces and create original music. Yoko, who directed and co-wrote the original game, collaborated with Masuyama on adapting the story into a form that would work in an anime.

Set in the far future during a proxy war between alien-created Machine Lifeforms and human-crafted androids, the story primarily focuses on 2B and 9S, soldiers for the elite YoRHa combat force who become involved with the latest conflict against the Machine Lifeforms. Based on the 2017 action role-playing game Nier: Automata developed by PlatinumGames and published by Square Enix, the anime is being produced by A-1 Pictures. Nier: Automata Ver1.1a is a 2023 Japanese animated television series directed by Ryouji Masuyama, co-written by Masuyama and Yoko Taro, and composed by music studio Monaca.